I'm certainly no expert, but it seems like weapon reliability might have played a huge part in that. I was just reading the patrol history of the USS Tuna the other day, and it seemed to be filled with reports of firing fish only to have them run too deep or fail to detonate. I know how frustrating duds are in a computer simulation, I can only imagine how angry these crews must have gotten after hours (and sometimes days) of trying to set up a shot, only to have your main offensive weapon repeatedly fail on you.
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