You can run without a WO but if you try to plot or change course without a NA you'll stay on the last plotted course. Without a CE you can;t change speed, depth or course so its all to do with changes. The boat will go it's merry way without any officer in the control room but as soon as you want to change course, speed or depth then you've gotta have someone in both NA and CE positions. Interestingly the WO can be out of there for torpedo attacks as long as you are not requesting any info from him.
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