Before I put in my 2-cents worth, My German extends to a pitiful college semester on the subject and reading medieval "fechtbuchs" on German swordplay. Any native German speaker is more than welcome to correct me.
"los" in German is similar to the English verb, "go".
"Rohr Eins, los!"="Tube one, go!"
An example of this context comes (surprisingly enough) from the German dub of Spaceballs. Dark Helmet orders "Wahnsinnige Geschwindigkeit und los!" just before the iconic scene. Anyone who has seen the movie in English knows that the line is "Ludicrous speed, go!" The translation seems pretty direct.
The "fire at will" order, already stated as "Feuer Frei" literally translates into "fire free", which has equally obvious meaning.
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