Depends.
For submerged attacks, the commander would be at the periscope, calling out the relevant numbers (range, speed, AOB, running depth for eels, etc.) and those would be fed into the system.
For surface attacks, typically the 1WO (first officer, US equiv would be XO) would be at the UZO focusing on the targets and calling out that stuff while the commander remained in charge of the overall tactical situation.
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