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Make sure your CPU is properly seated in its socket.
Make sure the heat sink is properly seated on the CPU 
Make sure all the RAM is properly seated and a matched set.
Make sure your power supply is adaquate.
My money is on number 2
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I have used quality heat transfer paste for the heat sink plus I have software that checks the temperature and all is ok, once the computer gets that first beep will run without a hitch all day, the problem can be while the computer is still cold.

Power supply, maybe, ram is seated properly & always buy matched sets (2x2gb Kingston).
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I generally begin trouble shooting a cranky PC, by connecting the power supply to a test unit which gives you a visual representation of all the voltages on the different rails and other pertinent diagnostics. Once the PSU is considered healthy, then you can move on to checking out the MoBo, Memory and CPU in that order. The beeps on post are significant and if you go into the MoBo manual there is a chapter there on diagnosing and interpreting the beeps on post. You definately have a hardware issue, that is for sure.
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Would be good to have a test setup, but nothing spare here, I'm getting no post beeps it's either 1 beep or dead.
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Only advice I have is tell you what I would do: take it apart, completely. Clean all contacts/sockets/slots. Take your MB and PSU, connect the cables and power on and off untill you're sure it works. Add RAM, cycle power on and off, etc.
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This maybe the only way out!

Thing is
when the computer starts with the beep all runs ok, can leave it on overnight for a large download, works perfectly, and one thing I forgot to mention is that pressing the reset button works ok, starts without a hitch, well, has so far.

Times like these I need 2 computers.