If the game crashes to disk during mission loading but the training missions run OK that tends to indicate the 4GB patch (or the HSie hard coded fixes, which include it) is not installed - as suggested by the earlier posters. If the training missions do not run either then that says 3D graphics problems.
I'm afraid the best solution would be to rebuild the installation again or use one of the pre-cooked mod compilations like Onealex.
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